1. Crop Foundation Research Division, National Institute of Crop Science, Rural Development Administration, Wanju-gun 55365, Republic of Korea
2. Crop Protection Division, National Institute of Agricultural Sciences, Rural Development Administration, Wanju-gun 54875, Korea
3. Upland Crop Breeding Division, National Institute of Crop Science, Rural Development Administration, Miryang-si 50424, Korea
4. Division of Biotechnology, Jeonbuk National University, Jeonju-si 54896, Korea