1. Upland Flue-Cured Tobacco Quality and Ecology Key Laboratory of China Tobacco, Guizhou Academy of Tobacco Science, Guiyang 550081, P.R. China
2. College of Life Science, Yangtze University, Jingzhou 434025, P.R. China
3. Zheng’an County Tobacco Company, Zunyi Tobacco Company, Zunyi Zheng’an, Guizhou 563400, P.R. China
4. Key Laboratory of Molecular Genetics, Guizhou Academy of Tobacco Science, Guiyang 550081, P.R. China.