1. University of Reunion Island, UMR PVBMT, F-97410 St. Pierre, Reunion Island, France
2. CIRAD, UMR PVBMT, F-97410 St Pierre, Reunion Island, France
3. CIRAD, UMR PVBMT, 501 Toamasina, Madagascar
4. CIRAD, UMR AGAP, F-34398 Montpellier, France
5. AGAP, University of Montpellier, CIRAD, INRAE, Institut Agro, Montpellier, France