1. School of Life Science, Hunan University of Science and Technology, Xiangtan, Hunan 411201, China
2. Key Laboratory of Genetic Improvement and Multiple Utilization of Economic Crops in Hunan Province, Xiangtan, Hunan 411201, China
3. Key Laboratory of Ecological Remediation and Safe Utilization of Heavy Metal-polluted Soils, College of Hunan Province, Xiangtan, Hunan 411201, China
4. Institute of Crop Science, Chinese Academy of Agricultural Sciences, Beijing 100081, China
5. Key Laboratory of Integrated Management of the Pests and Diseases on Horticultural Crops in Hunan Province, Xiangtan, Hunan 411201, China