1. College of Food and Bioengineering, Henan University of Science and Technology, Luoyang 471003 Henan Province, China
2. University Key Laboratory of Microbiology Resources Development and Utilization, Henan University of Science and Technology, Luoyang 471023, China
3. College of Horticulture and Plant Protection, Henan University of Science and Technology, Luoyang 471003, China
4. International Peony Garden, Luoyang 471011, Henan Province, China
5. National Flower Park of China, Luoyang 471023, Henan Province, China
6. College of Agriculture/Tree Peony, Henan University of Science and Technology, Luoyang 471003, China
7. Henan Provincial Key Laboratory of Peony Characteristic Resource Plants, Henan University of Science and Technology, Luoyang 471003, Henan Province, China