1. First and second authors: Sydney Institute of Agriculture, School of Life and Environmental Sciences, University of Sydney, Biomedical Building C81, 1 Central Ave., Australian Technology Park, Eveleigh, NSW 2015, Australia; third author: Research School of Biology, College of Sciences, Australian National University, Canberra, ACT 2601, Australia; and fourth author: Plant Breeding Institute, School of Life and Environmental Sciences, University of Sydney, Private Bag 4011, Narellan, NSW 2567, Australia.