1. Department of Agronomy, Kansas State University, Manhattan, KS 66506, U.S.A.
2. National and Shandong Province Key Laboratories of Wheat Improvement, College of Agronomy, Shandong Agriculture University, Taian, Shandong 271018, China
3. College of Agronomy, Yangzhou University, Yangzhou, Jiangsu 225009, China
4. U.S. Department of Agriculture-Agricultural Research Service, Wheat, Peanut, and Other Field Crop Research Unit, Stillwater, OK 74075, U.S.A.
5. U.S. Department of Agriculture-Agricultural Research Service, Hard Winter Wheat Genetics Research Unit, Manhattan, KS 66506, U.S.A.