1. Hokkaido Agricultural Research Center, National Agriculture and Food Research Organization (HARC/NARO), Hitsujigaoka, Toyohira-ku, Sapporo, Hokkaido, 062-8555, Japan
2. Tokachi Agricultural Experiment Station, Hokkaido Research Organization, Memuro, Hokkaido 082-0081, Japan
3. Hokkaido-Chuo Station, Center for Seeds and Seedlings, NARO, Nishinosato, Kitahiroshima, Hokkaido 061-1102, Japan
4. Hokkaido Agricultural Research Center, National Agriculture and Food Research Organization (HARC/NARO), Hitsujigaoka, Toyohira-ku, Sapporo, Hokkaido, 062-8555, Japan.