1. Department of Plant Biology and Ecology, University of Antananarivo, Antananarivo 101, Madagascar
2. Centre Technique Horticole de Tamatave, Toamasina 501, Madagascar
3. CIRAD, UMR PVBMT, Saint Pierre 97410, La Réunion
4. CIRAD, UMR PVBMT, Toamasina 501, Madagascar