1. Sichuan University of Science and Engineering, College of Bioengineering, Zigong 643000, China
2. Liquor-Making Biotech and Application of Key Laboratory of Sichuan Province, Zigong 643000, China
3. State Key Laboratory of Mycology (SKLM), Institute of Microbiology, Chinese Academy of Sciences, Beijing 100101, China
4. Luzhou Laojiao Co. Ltd., Luzhou 646000, China