1. Holetta National Agricultural Biotechnology Research Center, Ethiopian Institute of Agricultural Research, Addis Ababa, Ethiopia
2. Biosciences east and central Africa-International Livestock Research Institute, Nairobi, Kenya
3. Centre for Tropical Crops and Biocommodities, Queensland University of Technology, Brisbane, Australia
4. Department of Biotechnology, Addis Ababa Science and Technology University, Addis Ababa, Ethiopia.