1. Stockbridge School of Agriculture, University of Massachusetts, Amherst, MA 01003
2. Department of Plant, Soil, and Microbial Sciences, Michigan State University, East Lansing, MI 48824
3. Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido 0606-0808, Japan
4. Stockbridge School of Agriculture, University of Massachusetts, Amherst, MA 01003; and Field Science Center for Northern Biosphere, Hokkaido University, Sapporo, Hokkaido 0606-0808, Japan