1. Key Laboratory of Horticultural Plant Biology, Huazhong Agricultural University, Wuhan 430070, Hubei, China, and Ministry of Education, China
2. Key Laboratory of Plant Pathology of Hubei Province, Huazhong Agricultural University, Wuhan 430070, Hubei, China
3. School of Environmental Sciences, University of Guelph, Guelph, ON N1G 2W1, Canada