1. INRAE, University of Bordeaux, BIOGECO, 33615, Pessac, France
2. INRAE, Bordeaux Sciences Agro, ISVV, SAVE, 33140, Villenave d'Ornon, France
3. Laboratory of Environmental Microbiology, Institute of Microbiology of the CAS, Vídeňská 1083, 14220 Praha 4, Czech Republic