1. Department of Virology, Wageningen Agricultural University, Binnenhaven 11, 6709 PD, Wageningen, The Netherlands, and Department of Crop Science, University of Zimbabwe, P.O. Box MP 167, Mount Pleasant, Harare, Zimbabwe
2. Department of Crop Science, University of Zimbabwe, P.O. Box MP 167, Mount Pleasant, Harare, Zimbabwe
3. Department of Virology, Wageningen Agricultural University, Binnenhaven 11, 6709 PD, Wageningen, The Netherlands