1. Department of Plant Pathology, Washington State University, Pullman, WA 99164-6430, U.S.A.
2. College of Life Science and Biotechnology, Mianyang Normal University, Mianyang, Sichuan 621010, China
3. Wheat Health, Genetics, and Quality Research Unit, U.S. Department of Agriculture–Agricultural Research Service, Pullman, WA 99164-6430, U.S.A.