1. CSIRO Plant Industry, Grain Quality Research Laboratory, PO Box 7, North Ryde, NSW 1670, Australia; and Quality Wheat Cooperative Research Center Ltd., Locked Bag 1345, North Ryde, NSW 1670, Australia.
2. Current address: School of Wine and Food Sciences, Charles Sturt University, Locked Bag 588, Wagga Wagga NSW 2678 Australia.
3. Corresponding author. Phone: +61 2 9490 8437. Fax: +61 2 9490 8419. E-mail:f.bekes@pi.csiro.au