1. Laboratory for Plant Virology, Department of Plant Pathology, College of Plant Protection, Shandong Agricultural University, Tai’an, Shandong 271018, China; and Shandong Provincial Key Laboratory of Agricultural Microbiology, Tai’an, Shandong 271018, China
2. Laboratory for Plant Virology, Department of Plant Pathology, College of Plant Protection, Shandong Agricultural University, Tai’an, Shandong 271018, China
3. Jiyang Plant Protection Station, Ji’nan, Shandong 251400, China
4. Laboratory for Plant Virology, Department of Plant Pathology, College of Plant Protection, Shandong Agricultural University, Tai’an, Shandong 271018, China; and Shandong Provincial Key Laboratory of Agricultural Microbiology, Tai’an, Shandong 271018, China.