1. Jeollabuk-do Agricultural Research and Extension Services, Iksan 54591, Korea
2. Department of Plant Science, Gangneung-Wonju National University, Gangneung 25457, Korea
3. Horticultural & Herbal Crop Environment Division, National Institute of Horticultural and Herbal Science, Wanju 55365, Korea
4. Division of Environmental Science and Ecological Engineering, Korea University, Seoul 02841, Korea.