1. Department of Agricultural Biotechnology, College of Agriculture and Life Sciences, Seoul National University, Seoul, 151-921, Republic of Korea
2. Center for Plant Aging Research, Institute for Basic Science (IBS), Daegu 42988, Republic of Korea
3. The Taejin Genome Institute, Gadam-gil 61, Hoeongseong, 25239, Republic of Korea
4. College of Crop Protection and Agronomy, Qingdao Agricultural University, Qingdao, Shandong 266109, China
5. Department of Agricultural Biotechnology, College of Agriculture and Life Sciences, Seoul National University, Seoul, 151-921, Republic of Korea, and College of Crop Protection and Agronomy, Qingdao Agricultural University, Qingdao, Shandong 266109, China.