1. Biosystem and Bioengineering Program, University of Science and Technology (UST), Daejeon 34141, Republic of Korea
2. Plant System Engineering Research Center, Korean Research Institute of Bioscience and Biotechnology, Daejeon 34141, Republic of Korea
3. Plant Biotechnology Research Division, Amhara Agricultural Research Institute, Bahir Dar, Ethiopia
4. Department of Plant Quarantine, Animal and Plant Quarantine Agency, Gimcheon 39660, Republic of Korea