1. Hainan Key Laboratory for Sustainable Utilization of Tropical Bioresource, College of Tropical Crops, Hainan University, Haikou 570228, Hainan, China
2. Jiaxing Academy of Agricultural Science, Jiaxing 314016, Zhejiang, China
3. College of Life Science, Hainan University, Haikou 570228, Hainan, China