1. College of Plant Science and Technology, Huazhong Agricultural University, Wuhan 430070, China
2. Institute of Chinese Herbal Medicines, Hubei Academy of Agricultural Sciences, Enshi 445000, China
3. Key Laboratory of Integrated Management of Crops of Central China, Ministry of Agriculture, P. R. China/Hubei Key Laboratory of Crop Disease, Insect Pests and Weeds Control, Hubei Academy of Agricultural Sciences, Wuhan 430064, China