1. Department of Biology, Texas State University, San Marcos, TX, U.S.A.;
2. Boyce Thompson Institute for Plant Research, Ithaca, NY, U.S.A.;
3. Department of Molecular Biosciences, The University of Texas at Austin, U.S.A.; and
4. USDA Robert W. Holley Center for Agriculture and Health, Tower Road, Ithaca, NY, U.S.A.