1. Institute of Biotechnology, Addis Ababa University, P.O. Box 1176, Addis Ababa, Ethiopia
2. Department of Biotechnology, Addis Ababa Science and Technology University, Addis Ababa, Ethiopia
3. Institute of Biotechnology, Addis Ababa University, Addis Ababa, Ethiopia, and Ethiopian Biotechnology Institute, Ministry of Science and Technology, Addis Ababa, Ethiopia
4. Leibniz Institute DSMZ-German Collection of Microorganisms and Cell Cultures, Inhoffenstrasse 7B, 38124 Braunschweig, Germany.