1. Key Laboratory of Integrated Pest Management on Tropical Crops, Ministry of Agriculture, Environment and Plant Protection Institute, Chinese Academy of Tropical Agricultural Sciences, Haikou 571101, China
2. Analysis and Test Center, Chinese Academy of Tropical Agricultural Sciences, Haikou, 571101, China
3. College of Plant Protection, Hainan University, Haikou 570228 China
4. Rothamsted Research, Harpenden, AL5 2JQ, U.K.
5. NIAB EMR, New Road, East Malling, Kent ME19 6BJ, U.K.