1. Department of Genetics and Plant Production; Aula Dei Experimental Station, EEAD-CSIC; Avda Montañana 1005 50059 Zaragoza Spain
2. The James Hutton Institute; Invergowrie DD2 5NE, Dundee Scotland UK
3. Institute for Crop Science and Plant Breeding; Bavarian State Research Centre for Agriculture; Am Gereuth 6 85354 Freising-Weihenstephan Germany