1. Department of Aquatic Bioscience Graduate School of Agricultural and Life Sciences University of Tokyo Tokyo Japan
2. Now at Yaeyama Field Station Fisheries Technology Institute Japan Fisheries Research and Education Agency Okinawa Japan
3. Now at Atmosphere and Ocean Research Institute University of Tokyo Chiba Japan
4. Now at Fisheries Resources Institute Japan Fisheries Research and Education Agency Yokohama Japan
5. Department of Ocean Sciences Tokyo University of Marine Science and Technology Tokyo Japan
6. Now at Graduate School of Fisheries and Environmental Sciences Nagasaki University Nagasaki Japan
7. Now at Graduate School of Science and Engineering Soka University Tokyo Japan