1. State Key Laboratory of Tropical Oceanography & Key Laboratory of Science and Technology on Operational Oceanography South China Sea Institute of Oceanology & Innovation Academy of South China Sea Ecology and Environmental Engineering Chinese Academy of Sciences Guangzhou China
2. University of Chinese Academy of Sciences Beijing China
3. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou China
4. Polar and Marine Research Institute, Jimei University Xiamen China