1. Flathead Lake Biological Station, Division of Biological Sciences,; University of Montana,; Polson, Montana, USA
2. Numerical Terradynamic Simulation Group, College of Forestry and Conservation,; University of Montana,; Missoula, Montana, USA
3. Climate Impacts Group, Joint Institute for the Study of the Atmosphere and Ocean and School of Aquatic and Fisheries Sciences,; University of Washington,; Seattle, Washington, USA