1. Key Laboratory of Alpine Ecology, Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China
2. Key Laboratory of Alpine Ecology, CAS Center for Excellence in Tibetan Plateau Earth Sciences and Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China
3. University of Chinese Academy of Sciences Beijing China
4. Key Laboratory of Mountain Surface Processes and Ecological Regulation of Chinese Academy of Sciences, Institute of Mountain Hazards and Environment Chinese Academy of Sciences Chengdu China
5. Key Laboratory of Petroleum Resources Research, Institute of Geology and Geophysics Chinese Academy of Sciences Lanzhou China
6. Department of Microbiology and Plant Biology and School of Civil Engineering and Environmental Sciences, Institute for Environmental Genomics University of Oklahoma Norman OK USA
7. Department of Environmental Health Sciences, School of Public Health University of Michigan Ann Arbor MI USA