1. Institute of Marine Science and Technology Shandong University Qingdao China
2. Research Centre for Indian Ocean Ecosystem Tianjin University of Science and Technology Tianjin China
3. College of Marine Science and Technology China University of Geosciences (Wuhan) Wuhan Hubei China
4. State Key Laboratory of Tropical Oceanography (LTO) South China Sea Institute of Oceanology Chinese Academy of Sciences Guangzhou China