1. CNRS, Univ Brest, IRD, Ifremer, LEMAR Plouzané France
2. Now at Department of Chemical Oceanography, Atmosphere and Ocean Research Institute University of Tokyo Tokyo Japan
3. EcoLab, Univ Toulouse, CNRS, INPT, UPS Toulouse France
4. LOG, UMR 8187 CNRS, Univ Lille, ULCO, Univ du Littoral Côte d'Opale Wimereux France
5. LEEISA, USR 3456 CNRS, Univ Guyane, Ifremer, Centre de recherche de Montabo Guyane Française France
6. Department of Biogeochemical Processes Max Planck Institute for Biogeochemistry Jena Germany