1. Faculty of Fisheries Sciences Hokkaido University Hakodate Japan
2. Arctic Research Center Hokkaido University Sapporo Japan
3. Field Science Center for Northern Biosphere Hokkaido University Hakodate Japan
4. Institute of Low Temperature Science Hokkaido University Sapporo Japan
5. National Institute of Polar Research Tachikawa Japan
6. Graduate University for Advanced Studies (SOKENDAI) Tachikawa Japan