1. Research Institute for Global Change Japan Agency for Marine‐Earth Science and Technology Yokosuka Japan
2. Department of Environmental Science and Technology, Graduate School of Bioresources Mie University Mie Japan
3. International Pacific Research Center, School of Ocean and Earth Science and Technology University of Hawaii Honolulu USA
4. Graduate School of Marine Science and Technology Tokyo University of Marine Science and Technology Tokyo Japan
5. College of Oceanography Hohai University Nanjing China