1. State Key Laboratory of Tropical Oceanography South China Sea Institute of Oceanology, Chinese Academy of Sciences Guangzhou China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou China
3. Institution of South China Sea Ecology and Environmental Engineering, Chinese Academy of Sciences Guangzhou China
4. CAS Key Laboratory of Tropical Marine Bio‐resources and Ecology South China Sea Institute of Oceanology, Chinese Academy of Sciences Guangzhou China
5. Jiangsu Ocean University Lianyungang China