1. Institute of Low Temperature ScienceHokkaido University Sapporo Japan
2. Arctic Research CenterHokkaido University Sapporo Japan
3. Graduate School of Environmental ScienceHokkaido University Sapporo Japan
4. National Institute of Polar Research Tachikawa Japan
5. National Institute of TechnologyTomakomai College Tomakomai Japan
6. Fisheries Research DivisionCity of Mombetsu Mombetsu Japan
7. Faculty of Fisheries SciencesHokkaido University Hakodate Japan