1. State Key Laboratory of Humid Subtropical Mountain Ecology of Ministry of Science and Technology Fujian Normal University Fuzhou China
2. Sanming Experimental Forest Sanming China
3. Department of Forestry and Natural Resources University of Kentucky Lexington KY USA
4. Department of Life Science National Taiwan Normal University Taipei Taiwan
5. Department of Earth Sciences Indiana University‐Purdue University Indianapolis (IUPUI) Indianapolis IN USA
6. Department of Plant and Soil Sciences University of Kentucky Lexington KY USA
7. College of Mathematics and Statistics Hainan Normal University Haikao China