1. Graduate School of Agriculture; Kyoto University; Kyoto Japan
2. Center for Ecological Research; Kyoto University; Otsu Japan
3. CREST, Center for Ecological Research, Japan Science and Technology Agency; Kyoto University; Otsu Japan
4. Interdisciplinary Graduate School of Science and Engineering; Tokyo Institute of Technology; Yokohama Japan
5. Institute of Symbiotic Science and Technology; Tokyo University of Agriculture and Technology; Fuchu Japan