1. Atmosphere and Ocean Research Institute; The University of Tokyo; Chiba Japan
2. Department of Aquatic Bioscience, Graduate School of Agricultural and Life Sciences; The University of Tokyo; Tokyo Japan
3. Research and Development Center for Global Change; Japan Agency for Marine-Earth Science and Technology; Yokosuka Japan
4. Marine Biological Section, Department of Biology; University of Copenhagen; Helsingør Denmark
5. Now at Division of Life Science; The Hong Kong University of Science and Technology; Hong Kong China
6. Department of Ocean Sciences; Tokyo University of Marine Science and Technology; Tokyo Japan
7. Now at Japan Sea National Fisheries Research Institute; Japan Fisheries Research and Education Agency; Niigata Japan
8. Now at Research Center for Subtropical Fisheries, Seikai National Fisheries Research Institute; Japan Fisheries Research and Education Agency; Okinawa Japan
9. Graduate School of Fisheries and Environmental Sciences; Nagasaki University; Nagasaki Japan
10. Now at Graduate School of Engineering; Soka University; Tokyo Japan