1. School of Marine Sciences Sun Yat‐sen University Zhuhai Guangdong China
2. Southern Marine Science and Engineering Guangdong Laboratory (Zhuhai) Zhuhai Guangdong China
3. Guangdong Provincial Key Laboratory of Marine Resources and Coastal Engineering Guangzhou Guangdong China
4. Pearl River Estuary Marine Ecosystem Research Station Ministry of Education Zhuhai Guangdong China
5. Guangdong Marine Development Planning Research Center Guangzhou Guangdong China