1. State Key Laboratory of Marine Environmental Science Xiamen University Xiamen China
2. Key Laboratory of Marine Ecosystem and Biogeochemistry, State Oceanic Administration & Second Institute of Oceanography Ministry of Natural Resources Hangzhou, China
3. College of Ocean and Earth Sciences Xiamen University Xiamen China
4. College of the Environment and Ecology Xiamen University Xiamen China
5. Aix Marseille Université, CNRS/INSU, Université de Toulon, IRD, Mediterranean Institute of Oceanography (MIO) Marseille France