1. Coral Reefs Research Center of ChinaGuangxi University Nanning China
2. Guangxi Laboratory on the Study of Coral Reefs in the South China SeaGuangxi University Nanning China
3. School of Marine SciencesGuangxi University Nanning China
4. South China Sea Institute of OceanologyChinese Academy of Sciences Guangzhou China
5. College of Ocean EngineeringGuangdong Ocean University Zhanjiang China