1. Center for Marine Environmental Studies (CMES), Ehime University
2. Faculty of Agriculture, Ehime University
3. Present address; Laboratory of Neutraceuticals and Functional Food Science, Graduate School of Marine Science and Technology, Tokyo University of Marine Science and Technology
4. Integrated Center of Scienc+e (INCS), Ehime University
5. South Ehime Fisheries Research Center, Ehime University