1. Department of Cellular Signaling, Graduate School of Pharmaceutical Sciences, Tohoku University
2. Department of Physiology, Faculty of Medicine, Srinakharinwirot University
3. Institute for International Advanced Interdisciplinary Research, Tohoku University International Advanced Research and Education Organization
4. Department of Biological Informatics and Experimental Therapeutics, Akita University Graduate School of Medicine
5. Present address of corresponding author. Department of Biochemistry, Tokyo Women’s Medical University
6. Department of Natural Products Chemistry, Graduate School of Pharmaceutical Sciences, Tohoku University