1. All Out Africa Research Unit, Department of Biological Sciences, University of Swaziland, Private Bag 4, Kwaluseni, Swaziland
2. School of Natural Resources and Environment and Department of Wildlife Ecology and Conservation University of Florida, Newins-Ziegler Hall, Gainesville FL 32611, USA
3. Mammal Research Institute, Department of Zoology & Entomology, University of Pretoria, Private Bag 20, Hatfield 0028, Pretoria, South Africa
4. Centre for Biodiversity and Conservation Science, University of Queensland, Brisbane, Queensland 4072, Australia