1. Department of Biological Sciences, University of Eswatini, Private Bag 4, Kwaluseni, Eswatini
2. Department of Ecology and Evolutionary Biology, Princeton University, 106A Guyot Hall, Princeton, NJ 08544, USA
3. E.O. Wilson Biodiversity Laboratory, Gorongosa National Park, Mozambique
4. Durban Natural Science Museum, PO Box 4085, Durban 4000, South Africa
5. South African National Biodiversity Institute, P.O. Box 754, Pretoria, 0001, South Africa