1. Department of Evolutionary Ecology, Doñana Biological Station (CSIC), Av. Americo Vespucio 26, 41092 Sevilla, Spain
2. Department of Statistic and Operations Research, University of Seville, Faculty of Mathematics, Av. Reina Mercedes s/n, 41012 Sevilla, Spain
3. Department of Biology, Lund University, S-223 62 Lund, Sweden