1. State Key Laboratory of Biocontrol, Key Laboratory of Biodiversity Dynamics and Conservation of Guangdong Higher Education Institute, College of Ecology and Evolution, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China; e-mails: ,
2. Laboratory of Molecular Systematics, Department of Zoology, Faculty of Science, Palacky University, Olomouc, Czech Republic;