1. Key Laboratory of Biodiversity Dynamics and Conservation of Guangdong Higher Education Institute, the Museum of Biology, School of Life Sciences, Sun Yat-sen University, Guangzhou 510275, China;
2. College of Life Sciences, Capital Normal University, Xisanhuanbeilu 105, Haidian District, Beijing, 100048, P.R. China;
3. Guangzhou Institute of Forestry and Landscape Architechture, Guangzhou 510405, Guangdong, China
4. Department of Entomology, Moravian Museum, Brno, Czech Republic;
5. Australian National Insect Collection, CSIRO, National Research Collections Australia, GPO Box 1700, Canberra, ACT 2601, Australia;